These reprints may be used for your own personal educational/scientific use. Any further distribution, publication or commercial usage of this copyrighted material would require permission from the appropriate publisher.

** graduate or *** undergrad student advisee / mentee

Schneider**, R. S. and K. E. Boyer.  In press.  Limiting life history stages in the endangered wetland plant, Cirsium hydrophilum var. hydrophilum (Asteraceae). Madroño.

Burdick, D. M., G. E. Moore, and K. E. Boyer.  In press.  Impacts of exotic and native species invading tidal marshes. Chapter 14 in Salt Marshes, Z. Hughes and D. Fitzgerald, eds., Cambridge University Press.

Hovel, K. A., J. E. Duffy, J. J. Stachowicz, P. Reynolds, C. Boström, K. E. Boyer, M. Cusson, F. J. Fodrie, K. Gagnon, C. M. Hereu, M. Hori, P. Jorgensen, C. Kruschel, K. Lee, M. Nakaoka, N. E. O’Connor, F. Rossi, J. Ruesink, F. Tomas, and S. Ziegler. 2021. Joint effects of patch edges and habitat degradation on faunal predation risk in a widespread marine foundation species. Ecology Early view

Cloern, J. E., S. Safran, L. S. Vaughn, A. Robinson, A. A. Whipple, K. E. Boyer, J. Z. Drexler, R. J. Naiman, J. L. Pinckney, E. R. Howe, E. A. Canuel, and J. L. Grenier.  2021. Human appropriation of wetland primary production. Science of the Total Environment PDF

Hansen***, M. L., K. Crow, M. V. Buchbinder, and K. E. Boyer.  2020.  Invasive mangroves produce unsuitable habitat for endemic goby and burrowing shrimp pairs in Kāne'ohe Bay, Oahu, Hawai'i. Ciencias Marinas Western Society of Naturalists special issue. PDF Special issue collage

Grimes**, T. M., M. T. Tinker, B. B. Hughes, K. E. Boyer, L. Needles, K. Beheshti**, and R. L. Lewison.  2020. Characterizing the impact of recovering sea otters on commercially important crab in California estuaries. Marine Ecology Progress Series PDF

Hernán, G., M. J. Ortega, J. Henderson, J. Alos, K. Boyer, S. Cimon, V. Combes, M. Cusson, C. M. Hereu, M. Hessing-Lewis, K. Hovel, P. Jorgensen, S. Kiriakopolos, N. Kollars, M. I. O´Connor, J. Olsen, P. L. Reynolds, J. Ruesink, E. Voigt and F. Tomas.  2020.  Latitudinal variation in plant defense against herbivory in a marine foundation species does not follow a linear pattern: the importance of resource availability. Global Ecology and Biogeography. PDF

Rudebusch**, J., B. Hughes, K. E. Boyer, and E. Hines.  2020.  Assessing anthropogenic risk to sea otters (Enhydra lutris nereis) for reintroduction into San Francisco Bay. PeerJ PDF

Parker, V. T and K. E. Boyer.  2019.  Sea-level rise and climate change impacts on an urbanized Pacific Coast Estuary. Wetlands 39:1219–1232. PDF

Hughes, B. B., K. Wasson, T. Tinker, S. L. Williams, L. P. Carswell, K. E. Boyer, M. W. Beck, R. Eby, R. Scoles, M. Staedler, S. Espinosa, M. Hessing-Lewis, E. U. Rechsteiner, K. Beheshti, T. M. Grimes, B. H. Becker, L. Needles, J. A. Tomoleoni, J. Rudebusch, E. Hines, and B. R. Silliman. 2019. Species recovery and recolonization of past habitats: lessons for science and conservation from sea otters in estuaries. PeerJ 7:e8100 PDF

Wittyngham**, S. S., J. Moderan, and K. E. Boyer.  2019.  Temperature and salinity effects on submerged aquatic vegetation traits and susceptibility to grazing. Aquatic Botany 158: 103119. PDF

Harrison, C. D., et al. (scores of authors, including K. Boyer).  2019.  Investigating instructor talk in novel contexts: widespread use, unexpected categories, and an emergent sampling strategy. CBE: Life Sciences Education 18:ar47. PDF

Rohr, M. E. et al. (36 authors, including K. Boyer).  2018.  Blue carbon storage capacity of temperate eelgrass (Zostera marina) meadows. Global Biogeochemical Cycles 32:1457-1475. PDF, Supplemental Material PDF

Owens, M. T., et al., (scores of authors, including K. Boyer).  2018.  Collectively improving our teaching: Attempting biology department-wide professional development in scientific teaching. CBE: Life Sciences Education. PDF

Reynolds, P. L, J. J. Stachowicz, K. Hovel, C. Boström, K. Boyer, M. Cusson, J. S. Eklöf, F. G. Engel, A. H. Engelen, B. K. Eriksson, J. Fodrie, J. N. Griffin, C. Hereu, M. Hori, T. Hanley, M. Ivanov, P. Jorgensen, C. Kruschel, K.-S. Lee, K. McGlathery, P. O. Moksnes, M. Nakaoka, F. T. Nash,  M. I. O'Connor, N. O'Connor, R. J. Orth, F. Rossi, J. Ruesink, E. Sotka, R. K. F. Unsworth, M. A. Whalen, J. E. Duffy.  2018.  Latitude, temperature and habitat complexity predict predation pressure in eelgrass beds across the Northern Hemisphere. Ecology 99(1):29-35. PDF

Kollars, N. M., A. K. Henry, M. A. Whalen, K. E. Boyer, M. Cusson, J. S Eklöf, C. Hereu, P. Jorgensen, S. Kiriakopolos**, P. L. Reynolds, F. Tomas, M. S. Turner, and J. L. Ruesink.  2017.  Meta-analysis of reciprocal linkages between temperate seagrasses and waterfowl with implications for conservation. Frontiers in Plant Science. PDF

Murphy, J. L., K. E. Boyer, and E. C. Carpenter.  2017.  Restoration of cordgrass salt marshes: limited effects of organic matter additions on nitrogen fixation. Wetlands. PDF

Owens, M. T., et al. (scores of authors including K. Boyer).  2017.  Classroom sound can be used to classify teaching practices in college science courses. Proceedings of the National Academy of Sciences 114 (12) 3085-3090. PDF

Boyer, K., C. Zabin, S. De La Cruz, E. Grosholz, M. Orr, J. Lowe, M. Latta, J. Miller, S. Kiriakopolos**, C. Pinnell**, D. Kunz, J. Moderan, K. Stockmann**, G. Ayala, R. Abbott, and R. Obernolte.  2017.  San Francisco Bay Living Shorelines: Restoring Eelgrass and Olympia Oysters for Habitat and Shore Protection. Chapter 17 in D. M. Bilkovic, M. Mitchell, J. Toft, and M. La Peyre, eds., Living Shorelines: The Science and Management of Nature-Based Coastal Protection. CRC Press Marine Science Series. PDF

Cloern, J. E., A. Robinson, A. Richey, L. Grenier, R. L. Grossinger, K. E. Boyer, J. Burau, E. A. Canuel, J. F. DeGeorge, J. Z. Drexler, C. Enright, E. R. Howe, R. Kneib, A. Mueller-Solger, R. J. Naiman, A. Mueller-Solger, J. L. Pinckney, D. Schoellhamer, and C. Simenstad. 2016.  Primary production in the Delta, then and now.  San Francisco Estuary and Watershed SciencePDF

Robinson, A., A. Richey, J. E. Cloern, K. E. Boyer, J. Burau, E. Canuel, J. DeGeorge, J. Z. Drexler, L. Grenier, R. L. Grossinger, E. R. Howe, R. Kneib,  R. J. Naiman, A. Mueller-Solger, J. L. Pinckney, D. Schoellhamer, and C. Simenstad. 2016.  Primary Production in the Sacramento-San Joaquin Delta: A science strategy to quantify change and identify future potential. A Report of SFEI-ASC's Resilient  Landscapes Program, Publication #781, San Francisco Estuary Institute-Aquatic Science Center, Richmond, CA.  PDF

Boyer, K. E., and M. Sutula.  2015.  Factors controlling submersed and floating macrophytes in the Sacramento-San Joaquin Delta. Southern California Coastal Water Research Project. Technical Report No. 870. Costa Mesa, CA. PDF

Borgnis**, E. and K. E. Boyer.  2015.  Salinity tolerance and competition drive distributions of native and invasive submerged aquatic vegetation in the upper San Francisco Estuary.  Estuaries and Coasts DOI 10.1007/s12237-015-0033-5  PDF

Lewis**, J. T. and K. E. Boyer.  2014.  Grazer functional roles, induced defenses, and indirect interactions: Implications for eelgrass restoration in San Francisco Bay. Diversity 6: 751-770.  PDF

Archbald**, G. and K. E. Boyer.  2014.  Potential for spread of Algerian sea lavender (Limonium ramosissimum sbsp. provinciale) in tidal marshes.  Invasive Plant Science and Management 7:454-463.  PDF

Carr**, L. A. and K. E. Boyer.  2014.  Variation at multiple trophic levels mediates a novel seagrass-grazer interaction. Marine Ecology Progress Series 508:117–128.  PDF

Archbald**, G. and K. E. Boyer.  2014.  Distribution and invasion potential of Limonium ramosissimum subsp. provinciale in San Francisco Estuary salt marshes. San Francisco Estuary and Watershed Science 12(2)  PDF

Ort, B. S., C. S. Cohen, K. E. Boyer, L. K. Reynolds, S. M. Tam and S. Wyllie-Echeverria.  2014.  Conservation of eelgrass (Zostera marina) genetic diversity in a mesocosm-based restoration experiment.  PLoS ONE DOI: 10.1371/journal.pone.0089316.  PDF

Powers, S. P. and K. E. Boyer.  2014.  Marine Restoration Ecology.  Ch. 22 in M. D. Bertness, J. P. Bruno, B. R. Silliman, and J. J. Stachowicz, eds., Marine Community Ecology and Conservation.  Sinauer Associates.

Boyer, K. E. and W. J. Thornton**.  2012.  Natural and restored tidal marsh communities.  Chapter 17 in A. Palaima, ed., Ecology, Conservation, and Restoration of Tidal Marshes: The San Francisco Estuary.  University of CA Press.  PDF

Ort., B. S., C. S. Cohen, K. E. Boyer, and S. Wyllie-Echeverria.  2012.  Genetic diversity within and among eelgrass (Zostera marina) beds in the San Francisco Bay.  Journal of Heredity 103:533-546.  PDF

Ryan**, A. B. and K. E. Boyer.  2012.  Nitrogen further promotes a dominant salt marsh plant in an increasingly saline environment.  Journal of Plant Ecology 5:429-441.  PDF

Reynolds, L. K., L. A. Carr**, and K. E. Boyer.  2012.  Invasive amphipod (Ampithoe valida) consumption of eelgrass (Zostera marina) inflorescences in San Francisco Bay.  Marine Ecology Progress Series 451:107-118.  PDF

Carr**, L. A., K. E. Boyer, and A. Brooks.  2011.  Spatial patterns in epifaunal community structure in San Francisco Bay eelgrass (Zostera marina) beds.  Marine Ecology. 32: 88-103.  PDF

Boyer, K. E. and A. P. Burdick**.  2010.  Control of Lepidium latifolium (perennial pepperweed) and recovery of native plants in tidal marshes of the San Francisco Estuary.  Wetlands Ecology and Management 18:731-743.  PDF  ErrataCorrected Fig. 4, Corrected Table 2.

Boyer, K. E. and S. Wyllie-Echeverria.  2010.  Eelgrass Conservation and Restoration in San Francisco Bay: Opportunities and Constraints.  Report for the San Francisco Bay Subtidal Habitat Goals Project (Appendix 8-1).  PDF

Reynolds, L. K. and K. E. Boyer.  2010.  Perennial pepperweed (Lepidium latifolium): properties of invaded tidal marshes.  Invasive Plant Science and Management 3:130-138.  PDF

Boyer, K. E., J. S. Kertesz**, and J. F. Bruno.  2009.  Biodiversity effects on productivity and stability of marine macroalgal communities: the role of environmental context.  Oikos 118:1062-1072.  PDF


Bruno, J. F., K. E. Boyer, S. C. Lee, and J. E. Duffy.  2008.  Relative and interactive effects of plant and grazer richness in a benthic marine community. Ecology 89:2518-2528PDF

Huntington**, B. E. and K. E. Boyer.  2008.  Evaluating patterns of nitrogen supply using macroalgal tissue content and stable isotopic signatures in Tomales Bay, CA.  Environmental Bioindicators 3:180-192.  PDF

Huntington**, B. E. and K. E. Boyer.  2008.  Impacts of red macroalgal abundance (Gracilariopsis sp.) on eelgrass (Zostera marina) in Tomales Bay, California.  Marine Ecology Progress Series 267:133-142. PDF

Armitage, A. R., K. E. Boyer, R. R. Vance, and R. F. Ambrose.  2006.  Restoring assemblages of salt marsh halophytes in the presence of a rapidly colonizing dominant species. Wetlands 26:667-676.  PDF

Boyer, K. E., and P. Fong.   2005.  Co-occurrence of habitat-modifying invertebrates: effects on structural and functional properties of a created salt marsh.  Oecologia 143:619-628.  PDF

Boyer, K. E., and P. Fong.   2005.  Macroalgal-mediated transfers of water column nitrogen to intertidal sediments and salt marsh plants.  Journal of Experimental Marine Biology and Ecology 321:59-69.  PDF

Bruno, J. F., K. E. Boyer, J. E. Duffy, S. C. Lee, and J. S. Kertesz**. 2005.  Relative effects of species identity and richness on primary production in benthic marine communities.  Ecology Letters 8:1165-1174.  PDF

Boyer, K. E., P. Fong, A. R. Armitage, and R. A. Cohen.  2004.  Elevated nutrient content of macroalgae increases rates of herbivory in coral, seagrass, and mangrove habitats.  Coral Reefs 23:530-538.  PDF

Fong, P., K. E. Boyer, K. Kamer, and K. A. Boyle.  2003.  Influence of initial tissue nutrient status of tropical marine algae on response to nitrogen and phosphorus additions.  Marine Ecology Progress Series 262:111-123.  PDF

Lindig-Cisneros, R., J. Desmond, K. E. Boyer, and J. B. Zedler.  2003.  Wetland restoration thresholds: can a degradation transition be reversed with increased effort?  Ecological Applications 13:193-205.  PDF

Boyer, K. E., P. Fong, R. R. Vance, and R. F. Ambrose.  2001.  Salicornia virginica in a southern California saltmarsh: seasonal patterns and a nutrient enrichment experiment.  Wetlands 21:315-326.  PDF

Fong, P., K. Kamer, K. E. Boyer, and K. A. Boyle.  2001.  Nutrient content of macroalgae with differing morphologies may indicate sources of nutrients to tropical marine systems.  Marine Ecology Progress Series 220:137-152.  PDF

Boyer, K. E., J. C. Callaway, and J. B. Zedler.  2000.  Evaluating the progress of restored cordgrass (Spartina foliosa) marshes: belowground biomass and tissue N.  Estuaries 23:711-721.  PDF

Boyer, K. E. and J. B. Zedler.  1999.  Nitrogen addition could shift plant community composition in a restored California salt marsh.  Restoration Ecology 7:74-85.  PDF

Boyer, K. E. and J. B. Zedler.  1998.  Effects of nitrogen additions on the vertical structure of a constructed cordgrass marsh.  Ecological Applications 8:692-705.  PDF

Fong, P., K. E. Boyer, and J. B. Zedler.  1998.  Developing an indicator of nutrient enrichment in coastal estuaries and lagoons using tissue nitrogen content of the opportunistic alga, Enteromorpha intestinalis (Link).  Journal of Experimental Marine Biology and Ecology 231:63-79.  PDF

Zedler, J. B., K. E. Boyer, G. D. Williams, and J. C. Callaway.  1998.  Creating intertidal habitat for endangered species in southern California: achievements and challenges.  In A. McComb and J. Davis, eds.  Wetlands for the Future.  Gleneagles Publishing, Adelaide, Australia.

Haltiner, J., J. B. Zedler, K. E. Boyer, G. D. Williams, and J. Callaway.  1997.  Influence of physical processes on the design, functioning and evolution of restored tidal wetlands in California (USA).  Wetlands Ecology and Management 4:73-91. PDF

Zedler, J. B., J. Desmond, S. Phinn, B. Nyden, G. Sullivan, G. D. Williams, J. C. Callaway, K. E. Boyer, and A. Powell.  1997.  New tools for assessing coastal habitats.  Proceedings of the 1997 California and the World Oceans meeting, San Diego, California.

Boyer, K. E. and J. B. Zedler.  1996.  Damage to cordgrass by scale insects in a constructed salt marsh: effects of nitrogen additions.  Estuaries 19:1-12.  PDF

Fong, P., K. E. Boyer, J. S. Desmond, and J. B. Zedler.  1996.  Salinity stress, nitrogen competition, and facilitation: what controls seasonal succession of two opportunistic green macroalgae?  Journal of Experimental Marine Biology and Ecology 206:203-221.  PDF


A. Ryan