Lecturer and Adjunct Associate Professor, Department of Biology,

Senior Research Scientist, Romberg Tiburon Centers

San Francisco State University, San Francisco, CA

1981-82 Visiting Postdoctoral Fellow, University of California, Los Angeles, CA, (occasional lecturing)

1984-89 Research Associate, Allan Hancock Foundation, University of Southern California (occasional lecturing)

1989- Assistant Research Professor, Department of Biological Sciences, University of Southern California

1994 Visiting Scientist, Institute of Marine Biology, Crete, Greece

1996 Acting Director, Graduate Program in Ocean Sciences, University of Southern California

1996 Lecturer, Adjunct Faculty, Departmnet of Biology, San Francisco State University, San Francisco, CA

1996 Associate Research Faculty, Department of Biological Sciences, University of Southern California

Wilkerson, P.2

1981 Pauley Foundation Grant-In-Aid to conduct research at Hawaii Institute of Marine Biology

Editor, Advances in Space Research

Journal Reviewer for Limnology & Oceanography,

Journal Reviewer for Biological Bulletin,

Journal Reviewer for Marine Biology,

Journal Reviewer for Marine Ecology Progress Series,

Journal Reviewer for Coral Reefs

Journal Reviewer for Journal of Exp. Marine Biology and Ecology

Proposal Reviewer for NSF Biological Oceanography,

Proposal Reviewer for NSF Chemical Oceanography, JGOFS

Proposal Reviewer for NSF Polar Programs,

Proposal Reviewer for SeaGrant, NOAA

Certified scuba diver (NAUI),

French, German

1. Wilkerson, F.P. 1980. Phosphorus in the green hydra symbiosis. Ph.D. Thesis, University of Bristol, UK. 164 p.

* 2. Wilkerson, F.P. 1980. Bacterial symbionts on green hydra and their effect on phosphate uptake. Microbial Ecology 6: 85-92.

* 3. Wilkerson, F.P. The phosphorus economy of green hydra symbiosis. In: Nutrition in the Lower Metazoa. D.C. Smith and Y. Tiffon (Eds). Pergamon Press. pp. 149-156.

4. Wilkerson, F.P. Symbionts involved in phosphate uptake by green hydra. 1980. In: Endocytobiology, Endosymbiosis and Cell Biology. Vol. 1. W. Schwemmler and H.E.A. Schenk (Eds). pp. 269-277.

* 5. Wilkerson, F.P., G. Muller Parker and L. Muscatine. 1983. Temporal patterns of cell division in natural populations of endosymbiotic algae. Limnology and Oceanography 28: 1009-1014.

* 6. Wilkerson, F.P. and L. Muscatine. 1984. Uptake and assimilation of dissolved inorganic nitrogen by a tropical sea anemone. Proceedings of the Royal Society, London. B 221: 71-86.

7. Wilkerson, F.P. and R.C. Dugdale. 1984. Possible connections between sewage effluent, nutrient levels and jelly-fish blooms. In: Proceedings of the workshop on jelly- fish blooms in the Mediterranean Sea, UNEP. pp. 195-201.

* 8. Wilkerson, F.P. and R.K. Trench. 1985. Nitrate assimilation by zooxanthellae. Marine Chemistry 16: 385-393.

9. Dugdale, R.C. and F.P. Wilkerson. 1985. Primary production in the Cap Blanc region. In: International Symposiun on Upwelling of West Africa., Inst. Inv. Pesq. C. Bas, R. Margalef and P. Rubies (Eds.), Barcelona 1: 233-243.

10. Dugdale, R.C. and F.P. Wilkerson.1985. Athens Project: far-field imact of surface versus deep sewage outfall, Woodward Clyde Associates Report 1, 28pp.

11. Dugdale, R.C. and F.P. Wilkerson. 1985. The design of the outfall diffuser system for the effluent from the treatment works on Psytallia Island and its diffusion in the Saronikos Gulf: Analysis of far-field impact, Woodward Clyde Associates Report 2, 50pp.

* 12. Dugdale, R.C. and F.P. Wilkerson. 1986. The use of 15N to measure nitrogen uptake in eutrophic oceans; experimental considerations. Limnology and Oceanography 31: 673-689.

* 13. Muscatine, L., F.P. Wilkerson and L.R. McCloskey. 1986. Regulation of population density of symbiotic algae in a tropical marine jellyfish (Mastigias sp.). Marine Ecology Progress Series 32: 279-290.

* 14. Wilkerson, F.P. and R.K. Trench. 1986. Uptake of dissolved inorganic nitrogen by the symbiotic clam Tridacna gigas and the coral Acropora sp. Marine Biology 93: 237-246.

* 15. Wilkerson, F.P. and R.C. Dugdale. 1987. The use of large shipboard barrels and drifters to study the effects of coastal upwelling on phytoplankton nutrient dynamics. Limnology and Oceanography 32: 368-382.

* 16. Wilkerson, F.P., R.C. Dugdale and R.T. Barber. 1987. Effects of El Nino on new, regenerated and total production in eastern boundary upwelling systems. Journal of Geophysical Research 92: 14347-14355.

* 17. Dugdale, R.C., F.P. Wilkerson, R.T. Barber, D. Blasco and T.T. Packard. 1987. Changes in nutrients, pH, light penetration and heat budget by migrating photosynthetic organisms. Oceanology Acta Spec. 6: 103-109.

* 18. Wilkerson, F.P., D. Kobayashi and L. Muscatine. 1988. Cell divison and size of symbiotic algae in Caribbean reef corals. Coral Reefs 7: 29-36.

* 19. Dugdale, R.C. and F.P. Wilkerson. 1988. Nutrient sources and primary production in the Eastern Mediterranean. Oceanologia Acta Spec: 179-184.

* 20. Dugdale, R.C. and F.P. Wilkerson. 1989. New production in the upwelling center at Point Conception, California: temporal and spatial patterns. Deep-Sea Research, 36: 985-1007.

* 21. Dugdale, R.C., A. Morel, A. Bricaud and F.P. Wilkerson. 1989. Modeling new production in upwelling centers: II A case-study of modeling new production from remotely sensed temperature and color. Journal of Geophysical Research, 94: 18119-18133.

22. Dugdale, R.C. and F.P. Wilkerson. 1989. Regional perspectives in global new production. In: Oceanologie actualite et propsective, M.M. Denis (ed.), Proceedings of the Centennial of Station d'Endoume, Marseille, pp. 289-309.

* 23. Dugdale, R.C., F.P. Wilkerson and A. Morel. 1990. Modeling new production in upwelling centers: I Applications of a shift-up model to 3 upwelling areas. Limnology and Oceanography, 35: 822-829

* 24. Dugdale, R.C. and F.P. Wilkerson. 1990. Iron addition experiments in the Antarctic- a re-analysis. Global Biogeochemical Cycles. 4:13-19.

* 25. Wilkerson, F.P. and G. Grunseich. 1990. Formation of blooms by the symbiotic ciliate Mesodinium rubrum: the significance of nitrogen uptake. Journal Plankton Research . 12: 973-989.

* 26. Dugdale, R.C. and F.P. Wilkerson. 1991. Low specific nitrate uptake rate: a common feature of high-nutrient, low-chlorophyll marine ecosystems. Limnology and Oceanography, 36: 1678-1689.

* 27. Dugdale, R.C., F.P. Wilkerson, R.T. Barber and F.P. Chavez. 1992. Estimating new production in the equatorial Pacific Ocean at 150oW. Journal Geophysical Research. 97: 681-686.

* 28. Wilkerson, F.P. and R.C. Dugdale. 1992 Measurements of nitrogen productivity in the equatorial Pacific at 150oW. Journal Geophysical Research. 97: 669-679.

* 29. Dugdale, R.C. and F.P. Wilkerson. 1992. Nutrient limitation of new production. In: Primary Productivity and Biogeochemical Cycles in the Sea, P.G. Falkowski and A.D. Woodhead (eds.), Plenum Press, New York, pp. 107-122.

* 30. Wilkerson, F.P. and P. Kremer. 1992. DIN, DON and PO4 uptake by a medusae with algal symbionts. Marine Ecology Prog. Ser. 90: 237-250

*31 Minas, H.J., R.C. Dugdale, M. Minas and F.P. Wilkerson. 1993. New production in oligotrophic waters: a review of results and ideas induced by the "14C-method" controversy.

* 32. Dugdale, R.C., F.P. Wilkerson, D.Halpern, F.P. Chavez and R.T. Barber. 1994. Remote sensing of seasonal and annual variation of equatorial new production: a model for global estimates. Advances in Space Research. 14:169-178.

* 33. McCloskey, L.R., L. Muscatine and F.P. Wilkerson. 1994. Daily photosynthesis, respiration, and carbon budgets in a tropical marine jellyfish (Mastigias sp.). Marine Biology 119: 13-22.

* 34. Dugdale, R.C., F.P. Wilkerson and H.J. Minas. 1994 The role of a silicate pump in driving new production. Deep-Sea Research 42: 697-719.

*35. Wilkerson, F.P. and R.C. Dugdale. Silicate versus nitrate limitation in the equatorial Pacific estimated from satellite-derived sea surface temperatures. Advances in Space Research. 18: 81-89.

*36. Raschke, E., V.V. Salomonson, R. Stuhlmann and F. Wilkerson. 1996. Satellite Data for Atmosphere, Continent and Ocean Research, p. 142, Elsevier Press.

*37. Dugdale, R.C., C.O. Davis and F.P. Wilkerson. 1997. Assessment of new production in the upwelling center at Point Conception, California estimated using remotely sensed sea-surface temperatures. Journal of Geophysical Research. 102: 573-8585

38. Wilkerson, F.P. and R.C. Dugdale. 1997. The effect of El Nino on coastal new production. In "California and the World Ocean ‘97, pp. 1554-1562.

*39. Dugdale, R.C. and F.P. Wilkerson. 1998. Silicate regulation of new production in the eastern equatorial Pcific, Nature. 391: 270:273.

*40. Wilkerson, F.P., R.C. Dugdale, F P Chavez, and R M Kudela. Biomass and productivity in Monterey Bay, California: contribution of the larger autotrophs. Subm. to Deep-Sea Res.

Wilkerson, F.P., G. Muller Parker and L. Muscatine. Temporal patterns of cell division in natural populations of endosymbiotic algae. 1982. Eos 63: 945-946.

Wilkerson, F.P. and L. Muscatine. 1984 .Enzymes involved in the assimilation of DIN by symbiotic algae. Eos 64: 1075-1076.

Wilkerson, F.P. and R.C. Dugdale. 1985 The use of large shipboard containers during OPUS-1983 to observe and simulate the effects of upwelling on phytoplankton nitrogen dynamics. Eos 66: 1288.

Dugdale, R.C. and F.P. Wilkerson. 1985. Nitrogen dynamics at Point Conception, California upwelling center during the non-El Nino and El Nino years, 1981 and 1983. Eos 66: 1288.

Wilkerson, F.P. and R.C. Dugdale. 1986. Applying the concept of new and regenerated production to coral reef symbioses. Eos 67: 993.

Wilkerson, F.P., R.C. Dugdale and G. Grunseich. 1987. The effect of light on DIN and DON uptake and bloom formation by Mesodinium rubrum in the Peru upwelling ecosystem at 15oS. Eos 68: 1707.

Wilkerson, F.P. and R.C. Dugdale. 1988 Shift-up and acceleration of nitrogen uptake in eastern equatorial Pacific upwelling. Eos 69: 1104.

Dugdale, R.C. and F.P. Wilkerson. 1988. New production in the eastern equatorial Pacific upwelling measured with 15N. Eos 69: 1135.

Dugdale, R.C. and F.P. Wilkerson. 1990. Evidence for a critical nitrate concentration in upwelling systems: implications for operation of the biological pump in the CO2 cycle. Eos 71: 111

Wilkerson, F.P., R.C. Dugdale and C.D. Kopczak. 1990. Measurements of new and regenerated production in the water column over Fieberling Guyot. Eos 71: 173.

Kopczak, C.D., F.P. Wilkerson and R.C. Dugdale. 1990. Structure of the nutricline near Fieberling Guyot (32o25'N, 127o47'W) in the eastern North Pacific. Eos 71: 174.

Yang, S.R., R.C. Dugdale and F.P. Wilkerson. 1992. New production in the equatorial Pacific in El Nino and La Nina conditions. Eos 73:295.

Wilkerson, F.P. and R.C. dugdale. 1994. Can silicate versus nitrate limitation be evaluated from remotely-sensed sea-surface temperature? Eos 73: 57.

White, K.K., R.C. Dugdale and F.P. Wilkerson. 1994. Silicate control of silicate and nitrate uptake in Monterey Bay, California. Eos 73: 57.

Dugdale, R.C. and F.P. Wilkerson. 1994. The biological silicate pump: effects on new production. Eos 75: 57.

Wilkerson, F.P., R.C. Dugdale, R.M. Kudela, W. Cochlan and T. Hayden. 1996. Seasonal new production at an upwelling site in Monterey Bay, CA; the contribution of the larger-celled autotrophs. Eos 76: 158.

Chavez, F.P., G.E. Friederich, R.C. Dugdale and F.P. Wilkerson. 1996. A study of the role of biology in the air-sea flux of CO2 during the Spring 95 MBL Experiment off Monterey Bay, CA, Eos 76: 129.

Wilkerson P.7

Dugdale, R.C., G.E. Friederich, F.P. Chavez, T. Pennington, F.P. Wilkerson and T. Hayden. 1996. Spatial distribution of new production in Monterey Bay during CoOP95:implications for CO2 air-sea flux. Eos 76: 158

1978 Bacteria associated with hydra. Society of Experimental Biology Symposium, Reading, U.K.

1979 The surface bacteria of hydra. Coelenterate Society, Oxford, U.K.

1979 The phosphorus economy of the green hydra symbiosis. Nutrition in the Lower Metazoa. Caen, France.

1980 The phosphorus metabolism of green hydra. Coelenterate Society, London, U.K.

1980 Symbionts involved in phosphate uptake by green hydra. Endocytobiology I. Tubingen, Germany.

1982 Temporal patterns of cell division in natural populations of endosymbiotic algae, AGU/ASLO San Francisco, CA.

1983 Tropical marine lakes of Palau, USC Seminar Series, Los Angeles, CA. Invited Seminar Speaker

1983 Possible connections between sewage effluent, nutrient levels and jelly-fish blooms. Uinted Nations Environment Program Jellyfish Symposium, Athens, Greece. Invited Speaker

1984 Enzymes involved in the assimilation of DIN by symbiotic algae. AGU Ocean Sciences/ASLO, New Orleans, LA.

1984 Nitrogen metabolism of symbiotic cnidarians. West Coast Coelenterate Colloquium, Friday Harbor, WA.

1984 Growth of zooxanthellae in situ in a reef coral. ASLO, Vancouver, Canada.

1985 Significance of nitrate uptake in symbiotic clams and corals. ASLO, Minneapolis, MN.

1985 The impact of nutrients from domestic sewage in the Saronikos Gulf. Oceanography of the Mediterranean Sea: interdisciplinary aspects of the pelagic environment. Villefranche-sur-Mer, France.

1986 The use of large shipboard containers during OPUS-1983 to observe and simulate the effects of upwelling on phytoplankton nitrogen dynamics. AGU/ASLO New Orleans, LA.

1986 An ecological requirement for "surge" uptake in dinoflagellates. ASLO, Kingston, RI. Chairman of Symbiosis Special Session.

1986 A comparison of nitrogen dynamics observed during El Nino years at Point Conception, California and 15oS, Peru. AGU Chapman Conference on El Nino, Quayaquil, Ecuador.

1986 Applying the concept of new and regenerated production to coral reef symbioses. AGU/ASLO San Francisco.

1987 Biologie des cnidaires symbiotiques, Villefranche-sur-Mer, France.

1987 Changes in primary nutrient availability and sea level at Catalina Island, California during the 1983 El Nino. ASLO Madison, WI.

Wilkerson P.8

1988 The effect of light on DIN and DON uptake and bloom formation by Mesodinium rubrum in the Peru upwelling ecosystem at 15oS. AGU/ASLO Ocean Sciences, New Orleans, LA.

1988 Nitrogen flux in marine symbioses. Rosenthiel School of Marine and Atmospheric Sciences, Miami, FL. Invited seminar.

1988 Shift-up and acceleration of nitrogen uptake in eastern equatorial Pacific upwelling. AGU/ASLO San Francisco, CA.

1989 Eastern equatorial Pacific measurements of nitrogen productivity used 15N during the WEC88 cruise. The Oceanography Society, Monterey, CA.

1989 N-15 and nutrient measurements obtained during TOPO 89 cruise to Fieberling Guyot. Presentation at TOPO PI meeting, University of Washington, Seattle, WA.

1990 Measurements of new and regenerated production in the water column over Fieberling Guyot. AGU/ASLO, New Orleans, LA.

1990 Nitrogen flux in animals that harbor symbiotic algae. USC, Los Angeles, CA, Invited seminar.

1992 Dynamics of inorganic and organic nitrogen uptake by symbiotic algae in a nutrient limited system. 1992 ASLO Aquatic Sciences Mtg, Santa Fe. NM.

1992 New production in the equatorial Pacific in El Nino and La Nina conditions. AGU, San Francisco, Ca. AGU

1993 Spatial and temporal variability of new production in Monterey Bay, CA. The Oceanography Society Meeting, Seattle, WA. (Wilkerson, et al.)

1993 The role of silicate in global new production: a neglected nutrient. The Oceanography Society Meeting, Seattle, WA. (Dugdale, Wilkerson and Minas)

1993 Silicic acid and nitrate uptake interactions in a coastal upwelling system. The Oceanography Society Meeting, Seattle, WA. (White, Dugdale, and Wilkerson).

1994 Seasonal maps of nutrients and new production in Monterey Bay during 1992, Monterey Bay Symposium, Monterey

1994 Co-Organizer of Special Session: Silicate Limitation in Marine Systems, AGU Ocean Sciences, San Diego, CA.

1994 Can silicate versus nitrate limitation be evaluated from remotely-sensed sea surface temperature? AGU Ocean Sciences, San Diego, CA. (Wilkerson and Dugdale)

1994 Silicate control of silicate and nitrate uptake in Monterey Bay, CA., AGU Ocean Sciences, San Diego, CA. (White, Dugdale and Wilkerson)

1994 The biological silicate pump: effects on new production, AGU Ocean Sciences, San Diego, CA. (Dugdale and Wilkerson).

1994 Nitrogen budgets of cnidarians that possess endosymbiotic algae. Invited, ASLO, Miami, FL.

1994 Delineation of high nitrate, low silicate, low chlorophyll in offshore upwelled waters using satellite derived sea-surface temperatures, COSPAR, Frankfurt, Germany (Dugdale and Wilkerson)

Wilkerson P.9

1995 The importance of silicate in driving new production: implications for remote sensing models, University of Southampton and Rennell Center for oceanography, England, UK. (Wilkerson and Dugdale)

1996 Organizer, Session on Eastern Boundary Systems off the California Coast, AGU-ASLO Ocean Sciences Mtg., San Diego, February

1996 Seasonal new production at an upwelling site in Monterey Bay, CA; the contribution of the larger-celled autotrophs, AGU-ASLO Ocean Sciences Mtg., San Diego, February (Wilkerson, Dugdale, Kudela, Cochlan and Hayden)

1996 A study of the role of biology in the air-sea flux of CO2 during the Sprimg 95 MBL Experiment off Monterey Bay, CA, AGU-ASLO Ocean Sciences Mtg., San Diego, February Invited (Chavez, Friederich, Dugdale, Wilkerson)

1996 Spatial distribution of new production in Monterey Bay during CoOP95:implications for CO2 air-sea flux, AGU-ASLO Ocean Sciences Mtg., San Diego, February (Dugdale, Friederich, Chavez, Pennington, Wilkerson and Hayden)

1996 The importance of the silicate pump in global ocean flux, OPALEO Workshop, University of Brest, France, June 1996 (Dugdale, Wilkerson and Minas).Invited

1996 The nutrient conditions required for order of magnitude changes in paleo-new production, AGU San Francisco, December 1996, Invited

1997 The relative importance of nitrate, ammonium and urea in detrmining estimates of eutrophic and oligotrophic new production. ASLO Ocean Sciences, Santa Fe, February Invited (Wilkerson and Dugdale)

1997 Is nitrate uptake the best candidate for measuring new production? ASLO Ocean Sciences, Santa Fe, February Invited (Dugdale and Wilkerson)

1997 The effect and prediction of El Nino on the coastal biological production of California. California and the World Ocean ‘97, San Diego, March 1997 (Dugdale and Wilkerson)

1197 Effects of massive flooding in January 1997 on the biology and physics of San Francisco Bay and the Gulf of the Farallones. ERF, Providence October. Bollens, Collins, Dugdale, Wilkerson, Cordell, Cloe, Cloern, Murphee, Rago, Hayden, Avent and Franklin)

1998 Ratios of carbon to nitrogen uptake by the Monterey Bay phytoplankton assemblage. AGU-ASLO Ocean Sciences Mtg., San Diego, February (Wilkerson, Dugdale, Chavez, Kudela, Pennington)

1998 Effects of meridional nutrient asymmetry in equatorial undercurrent source water. AGU-ASLO Ocean Sciences Mtg., San Diego, February (Dugdale, Wilkerson, Barber).

1981 Hawaii Institute of Marine Biology, Kaneohe, Hawaii. Nitrogen flux of corals and anemones

1982 Micronesian Mariculture Demonstration Center, Palau, W. Caroline Islands. Biology of the symbiotic jellyfish Mastigias sp.

1983 Discovery Bay Marine Laboratory, Jamaica. Effect of depth on coral reef zooxanthellae

1983 Micronesian Mariculture Demonstration Center, Palau, W. Caroline Islands. Nitrogen flux of giant clams

1984 Saronikos Gulf, Greece. Biology of jellyfish, DON uptake and stable isotope composition

1984 R/V VELERO IV, Point Conception, California. Sept. 16-28. Mapping upwelling plumes using shipboard analyses of nitrate reductase activity.

1987 Station Zoologique, France. Exploratory experiments to use 15N to trace and estimate salp assimilation

1988 R/V WECOMA, Honolulu to Tahiti, Feb. 17-Mar 17. Measuring equatorial new and regenerated production using on-board mass spectrometry.

1988 R/V CALANUS, Bahamas, Apr. 29-May. Investigating light/nutrient relationships in the symbiotic jellyfish Linuche.

1989 R/V NEW HORIZON, off California coast: Fieberling Guyot. Sept. 8-22, Mapping nitrate, ammonium and N productivity around a seamount.

1983 Workshop on Jellyfish Blooms, United Nation Environment Program, Athens, Greece.

1983 Coral Reef Workshop, American Society of Zoologists, Philadelphia, PA.

1984 Organization of Persistent Upwelling Structures (OPUS), P.I. Workshop. Lake Arrowhead, CA.

1984 Histocompatibility and Symbiosis of Aquatic Organisms, Los Angeles, CA.

1985 SCOR Working Group 56 Symposium. Vertical motion in the equatorial upper ocean and its effects upon living resources and the atmosphere, Paris, France.

1988 Flow over Abrupt Topography workshop, Scripps, La Jolla, CA.